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		LUQMetadata76
		Frog abundance in fertilization plots
		
			
				Lawrence L.
				Woolbright
			
			Siena College
			Investigator
			
				Department of Biology
				Loudonville
				NY
				12211-1462
				USA
			
			(518)783-2440
			voice
			LWoolbright@siena.edu
		
		
			
				Lawrence L.
				Woolbright
			
			Siena College
			Investigator
			
				Department of Biology
				Loudonville
				NY
				12211-1462
				USA
			
			(518)783-2440
			voice
			LWoolbright@siena.edu
		
		english
		
			
				Population estimates and body sizes of E. coqui were collected on fertilization, litter removal, and control plots. 
			
		
		
			productivity
			population density
			leptodactylidae
			eleutherodactylus coqui
			demography
			ancura
			amphibia
			Assigned by Principal Investigator
		
		
			population dynamics
			LTER Core
		
		
			tabonuco
			MRCE
			population dynamics
			amphibians
			LEF LTER Data Sets
		
		
			Animal Population Dynamics
			LEF LTER Data Categories
		
		
			
				ADEQUATE ACKNOWLEDGEMENT
				
					The scientists, the LEF-LTER Program, and the USDA International Institute of Tropical Forestry (IITF) must receive adequate acknowledgement for the use of data by other scientists, and the LEF LTER program must receive two copies of any publication using that data. One copy will be placed at the IITF library and one copy will stay with the LEF LTER Data Manager. A suggested format for this acknowledgement will be distributed along with the requested data.
				
				
					Acknowledgements for NSF Support.
					
						The following paragraph should be included in the acknowledgements of all publications funded in any way by current and future LTER grants:
						This research was performed under grant #DEB-0218039 from the National Science Foundation to the Institute of Tropical Ecosystem Studies (IEET), University of Puerto Rico, and the International Institute of Tropical Forestry (IITFR) as part of the Long-Term Ecological Research Program in the Luquillo Experimental Forest. Additional support was provided by the Forest Service (U.S. Department of Agriculture) and the University of Puerto Rico.
					
				
			
		
		
			
				http://luq.lternet.edu/data/lterdb76/metadata/lterdb76.htm#DATA
			
		
		
			
				The Luquillo Experimental Forest (LEF) covers 11,231 hectares and reaches 1079 m above sea level. It has a warm and wet climate and includes a variety of tropical habitats and forest types and streams, and a high diversity of plants and animals. A key feature of these ecosystems is disturbance. Hurricanes, landslides, and human disturbance have shaped the forest, and research by LUQ has stimulated a new appreciation of the significance of large-scale disturbances in tropical forested ecosystems and the key role of plants and animals in shaping the response to these events. Hurricanes occurring one and 10 years after LUQ began in 1988 provided landscape-scale natural experiments which are still followed closely.
				
					-65.92
					-65.7
					18.35
					18.23
					
						20
						1060
						meter
					
				
			
		
		
			
				Lawrence L.
				Woolbright
			
			Siena College
			Investigator
			
				Department of Biology
				Loudonville
				NY
				12211-1462
				USA
			
			(518)783-2440
			voice
			LWoolbright@siena.edu
		
		
			Luquillo Experimental Forest LTER
			
				University of Puerto Rico, Rio Piedras
				P. O. Box 21910
				San Juan
				PR
				00931-1910
				USA
			
			 (787) 764 - 0000 (ext. 4940)
			(787) 772 -1481
			nbrokaw@lternet.edu
			http://luq.lternet.edu
		
		
			
				
					
						Mark-recapture. Adult coquies (725 mm SVL) were and measured with dial calipers. Each was toedipped, individually in some plots, by cohorts or others. Sex and reproductive condition were noted. Juveniles were counted.
					
				
			
		
		
			Frog Population Dynamics
			
				
					Lawrence L.
					Woolbright
				
				Professor
			
			
				
					El Verde Station, located in the tabonuco forest zone on the NW side of the Luquillo Experimental Forest (LEF), has hosted numerous scientific efforts since the 1960s. Thus, a great deal of information exists concerning anuran populations at the site over the past 30 years. Although past surveys did not use consistent techniques, they clearly document the general status of the anuran community of the area. I began working at El Verde in the 1970s and have collected data on the anuran community primarily in the 1980s and 1990s. Until recently, these surveys also lacked a consistent sampling protocol, although they clearly document declines in species richness in comparison to earlier records.  Most Puerto Rican Eleutherodactylus are terrestrial frogs that breed for prolonged periods of time in more or less continuous habitat. Because their life cycle lacks a free living larvae stage, reproductive behavior is not tied to bodies of water and they do not have the large aggregations typical of many aquatic breeders. For these reasons, assessing their population status requires examining fairly large areas of habitat. I began systematically sampling the anuran community on a 12 ha grid at the Bisley watersheds in 1989 and a second 16 ha grid at El Verde in 1993. These efforts have provided a comprehensive data set that can be used to evaluate future changes in the anuran community in this forest. 
				
			
			
				
					LTER
				
			
		
		
			
				uid=LUQ,o=lter,dc=ecoinformatics,dc=org
				all
			
			
				public
				read
			
		
		
			allBlocksdata.txt
			Fertilization
			
				allBlocksdata.txt
				
					
						1
						\n
						\n
						column
						
							,
						
					
				
				
					
						http://luq.lternet.edu/data/lterdb76/data/allBlocksdata.txt
					
				
			
			
				
					geoReference
				
				
					
						
							1993-06-13
						
						
							1993-06-27
						
					
				
			
			
				
					
						
							Mark-recapture. Adult coquies (725 mm SVL) were and measured with dial calipers. Each was toedipped, individually in some plots, by cohorts or others. Sex and reproductive condition were noted. Juveniles were counted.
						
					
				
				
					
						
							
								geoReference
							
							
								temporalReferenceallBlocksdata.txt
							
						
					
					
						
					
				
			
			
				
					MRCE Plot Block Number
					MRCEBLOCKPLOT
					MRCE fertilization plot plot identifier using the same nopnmenclature as the original MRCE proyect. Each contain the corresponding MRCE Proyect's data variable -Block-; a Roman number representing a 20x20 m square with 10x10 m inner measurement plots.
					
						
							
								
									
										BLOCK I 
										 block names I in the MRCE project
									
									
										 BLOCK II 
										 block names II in the MRCE project
									
									
										 BLOCK III 
										 block names III in the MRCE project. All records have a value in this field.
									
								
							
						
					
				
				
					File name
					FILENAME
					file name designation
					
						
							
								
									file name designation, Control, Litter Removal and Fertilization are the three values.
								
							
						
					
				
				
					Sex of frog
					SEX
					Sex of captured frog; It is left blank if the sex could not be determined.
					
						
							
								
									
										F 
										 female
									
									
										 M 
										 male
									
								
							
						
					
				
				
					Frog ID (Identification)
					FROGID
					This codes the reproductive status; pattern morph; and toc-clip of each animal marked. The codification is as follows: A1 2345 b where A represents the reproductive status: male - female; 1 represents a pattern morph ( 9- wild type mottled; 11-vertebral stripe; 16-dorsalatesal stripes; 5-spots; 6-interocular bar; 1-smooth monocolor; 2-5=toe numbers); the 4 digit represent the 4 feet (2-left hand; 3-right; 4-left foot; 5-each ranges from 1-4 where its left most toe and 4 is right most); b means thumbs;Blank-is an optional and determined series representing no thumbs clipperd; T- is an optional and determined series meaning a tumb was found clipped on left footO-is an optional and determined series meaning a thumb was found clipped on right foot.
					
						
							
								
									This codes the reproductive status; pattern morph; and toc-clip of each animal marked. The codification is as follows: A1 2345 b where A represents the reproductive status: male - female; 1 represents a pattern morph ( 9- wild type mottled; 11-vertebral stripe; 16-dorsalatesal stripes; 5-spots; 6-interocular bar; 1-smooth monocolor; 2-5=toe numbers); the 4 digit represent the 4 feet (2-left hand; 3-right; 4-left foot; 5-each ranges from 1-4 where its left most toe and 4 is right most); b means thumbs;Blank-is an optional and determined series representing no thumbs clipperd; T- is an optional and determined series meaning a tumb was found clipped on left footO-is an optional and determined series meaning a thumb was found clipped on right foot.
								
							
						
					
				
				
					
					DATE6/14/1993
					
					
						
							
								
									
								
							
						
					
				
				
					Recapture Data for Date 1
					DATE1
					This shows whether each animal was seen on each night with an optional addition number shows reproductive state. Recapture data gathered on06/14/1993 when blocknumber is Block Iand Treatment is Control; Recapture data gathered on 06/13/1993 when blocknumber is Block Iand Treatment is Litter Removal or Fertilization; Recapture data gathered on 06/7/1993 when blocknumber is Block II for all Treatment; Recapture data gathered on 06/22/1993 when blocknumber is Block III for all Treatment; When left blank, the frog wasnot captured. Note: a "c" was used in earlier data files for 1990.
					
						
							
								
									
										X1
										=frog was captured that night and sex uncertain
									
									
										X2
										 frog was captured that night with baggy throat if a male or eggs visible but small if female;
									
									
										X3
										frog was captured that nightand found calling if male orwithgrand large eggs visible if female
									
								
							
						
					
				
				
					Recapture Data for Date 2
					DATE2
					This shows whether each animal was seen on each night with an optional addition number shows reproductive state.Recapture data gathered on 06/17/1993 when blocknumber is Block Iand Treatment is Control; Recapture data gathered on 06/14/1993 when blocknumber is Block Iand Treatment is Litter Removal or Fertilization; Recapture data gathered on 06/8/1993 when blocknumber is Block II and Treatment is Control or Control New or Fertilization; Recapture data gathered on 06/9/1993 when blocknumber is Block II and Treatment is Litter Removal; Recapture data gathered on 06/23/1993 when blocknumber is Block III for all Treatment.
					
						
							
								
									This shows whether each animal was seen on each night with an optional addition number shows reproductive state. Recapture data gathered on06/14/1993 when blocknumber is Block Iand Treatment is Control; Recapture data gathered on 06/13/1993 when blocknumber is Block Iand Treatment is Litter Removal or Fertilization; Recapture data gathered on 06/7/1993 when blocknumber is Block II for all Treatment; Recapture data gathered on 06/22/1993 when blocknumber is Block III for all Treatment; When left blank, the frog wasnot captured. Note: a "c" was used in earlier data files for 1990.
								
							
						
					
				
				
					Recapture Data for Date 3
					DATE3
					This shows whether each animal was seen on each night with an optional addition number shows reproductive state.Recapture data gathered on 06/18/1993 when blocknumber is Block Iand Treatment is Control; Recapture data gathered on 06/17/1993 when blocknumber is Block Iand TREATMENT is Litter Removal or Fertilization; Recapture data gathered on 06/9/1993 when blocknumber is Block II and TREATMENT is Control or Control New; Recapture data gathered on 06/10/1993 when blocknumber is Block II and TREATMENT is Litter Removal or Fertilization; Recapture data gathered on 06/27/1993 when blocknumber is Block III for all Treatment.
					
						
							
								
									This shows whether each animal was seen on each night with an optional addition number shows reproductive state.Recapture data gathered on 06/18/1993 when blocknumber is Block Iand Treatment is Control; Recapture data gathered on 06/17/1993 when blocknumber is Block Iand TREATMENT is Litter Removal or Fertilization; Recapture data gathered on 06/9/1993 when blocknumber is Block II and TREATMENT is Control or Control New; Recapture data gathered on 06/10/1993 when blocknumber is Block II and TREATMENT is Litter Removal or Fertilization; Recapture data gathered on 06/27/1993 when blocknumber is Block III for all Treatment.
								
							
						
					
				
				
					Recapture data for day 4
					DATE4
					This shows whether each animal was seen on each night with an optional addition number shows reproductive state. Recapture data gathered on 06/20/1993 when blocknumber is Block Iand Treatment is Control; Recapture data gathered on 06/18/1993 when blocknumber is Block Iand Treatment is Litter Removal or Fertilization; Recapture data gathered on 06/10/1993 when blocknumber is Block II and Treatment is Control or Control New; Recapture data gathered on 06/18/1993 when blocknumber is Block II and Treatment is Litter Removal or Fertilization.
					
						
							
								
									This shows whether each animal was seen on each night with an optional addition number shows reproductive state. Recapture data gathered on 06/20/1993 when blocknumber is Block Iand Treatment is Control; Recapture data gathered on 06/18/1993 when blocknumber is Block Iand Treatment is Litter Removal or Fertilization; Recapture data gathered on 06/10/1993 when blocknumber is Block II and Treatment is Control or Control New; Recapture data gathered on 06/18/1993 when blocknumber is Block II and Treatment is Litter Removal or Fertilization.
								
							
						
					
				
				
					Recapture data for day 5
					DATE5
					This shows whether each animal was seen on each night with an optional addition number shows reproductive state. Recapture data gathered on 06/18/1993 when blocknumber is Block II and Treatment is Control or Control New.
					
						
							
								
									This shows whether each animal was seen on each night with an optional addition number shows reproductive state. Recapture data gathered on 06/18/1993 when blocknumber is Block II and Treatment is Control or Control New.
								
							
						
					
				
				
					Body size
					BODYSIZE
					The body size from tip of snout to posterior tip of urostyle, taken with frog pressed flat against a notebook. Instrument measures to 0.1 mm sizes recorded to 0.5 mm measurement repeatable to � 1.0 mm. Body sizes ranges from 25.0 (Arbitrary cut-off adult status) to largest posible sizes of 50mm +
					
						
							
								millimeter
							
							
								real
							
						
					
				
				
					Sub-Total indicator
					COMMENTS
					Indicates when values represents sub-totals
					
						
							
								
									Indicates when values represents sub-totals
								
							
						
					
				
				
					Data collectorsm or observers
					DATACOLLECTORS
					Observers; people that obtained the data
					
						
							
								
									Observers; people that obtained the data
								
							
						
					
				
			
		
	
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